Physaria vitulifera |
Physaria chambersii |
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roundtip twinpod |
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
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Habit | Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). |
Stems | several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm. |
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
Basal leaves | blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse). |
(petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
Cauline leaves | blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute). |
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
Racemes | congested, (elongated in fruit). |
congested. |
Flowers | sepals oblong, 6–8 mm; petals spatulate, to 10 mm. |
sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (usually curving upward, sigmoid), 6–10 mm. |
(divaricate, slightly sigmoid), 8–15 mm. |
Fruits | didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep); valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose; replum oblong, often constricted, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
Seeds | flattened. |
flattened. |
2n | = 8, 16. |
= 8, 10, 16, 24. |
Physaria vitulifera |
Physaria chambersii |
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Phenology | Flowering Apr–Jun. | Flowering Apr–Jul. |
Habitat | Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas |
Elevation | 1600-3000 m (5200-9800 ft) | 1500-3200 m (4900-10500 ft) |
Distribution |
CO
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AZ; CA; ID; NV; OR; UT
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 665. | FNA vol. 7, p. 631. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) |
Web links |