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roundtip twinpod

Calder's bladderpod

Habit Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center).
Stems

several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm.

simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm.

Basal leaves

blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse).

blade oblanceolate, 2–3 cm, margins entire.

Cauline leaves

blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute).

(sessile or proximal shortly petiolate);

blade narrowly oblanceolate, 0.5–1.5 cm, margins entire.

Racemes

congested, (elongated in fruit).

loose.

Flowers

sepals oblong, 6–8 mm;

petals spatulate, to 10 mm.

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide).

Fruiting pedicels

(usually curving upward, sigmoid), 6–10 mm.

(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout).

Fruits

didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose;

replum oblong, often constricted, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence);

replum as wide as or wider than fruit;

ovules 10–14 per ovary;

style 1–2 mm.

Seeds

flattened.

plump.

2n

= 8, 16.

= 20.

Physaria vitulifera

Physaria calderi

Phenology Flowering Apr–Jun. Flowering Jun–Aug.
Habitat Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale Dry rocky summits, limestone flats and slopes, alpine knolls
Elevation 1600-3000 m (5200-9800 ft) 600-1500 m (2000-4900 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; YT
[BONAP county map]
Discussion

Physaria calderi is known from the Ogilvie and Richardson mountains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 665. FNA vol. 7, p. 629.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella calderi, Lesquerella arctica subsp. calderi
Name authority Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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