Physaria vitulifera |
Physaria bellii |
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roundtip twinpod |
Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
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Habit | Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. |
Stems | several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm. |
simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
Basal leaves | blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse). |
(strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
Cauline leaves | blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute). |
blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
Racemes | congested, (elongated in fruit). |
dense. |
Flowers | sepals oblong, 6–8 mm; petals spatulate, to 10 mm. |
sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
Fruiting pedicels | (usually curving upward, sigmoid), 6–10 mm. |
(divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
Fruits | didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep); valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose; replum oblong, often constricted, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
Seeds | flattened. |
compressed. |
2n | = 8, 16. |
= 8. |
Physaria vitulifera |
Physaria bellii |
|
Phenology | Flowering Apr–Jun. | Flowering Mar–Jun(-Jul). |
Habitat | Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale | Dark shale, road cuts, ridge crests, washes |
Elevation | 1600-3000 m (5200-9800 ft) | 1500-1800 m (4900-5900 ft) |
Distribution |
CO
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CO
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Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 665. | FNA vol. 7, p. 628. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) |
Web links |