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roundtip twinpod

arctic bladderpod

Habit Perennials; caudex simple or branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate or bifurcate, (relatively massive, smooth to few-tubercled). Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth).
Stems

several from base, usually decumbent to ascending, (arising laterally, unbranched, coarse), 1–2 dm.

simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm.

Basal leaves

blade pandurate or obovate, 3–6 cm, margins usually deeply and broadly incised, rarely subentire, (apex obtuse).

(usually ± rosulate);

blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire.

Cauline leaves

blade oblanceolate to spatulate, similar to basal, (3–6 mm wide), margins entire, (apex often somewhat acute).

(sessile or shortly petiolate);

blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire.

Racemes

congested, (elongated in fruit).

loose.

Flowers

sepals oblong, 6–8 mm;

petals spatulate, to 10 mm.

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw).

Fruiting pedicels

(usually curving upward, sigmoid), 6–10 mm.

(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout).

Fruits

didymous, irregular in shape, somewhat angular, inflated, 5–7 × 6–8 mm, (papery, often rigid, base obtuse or truncate, apical sinus broad, open and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes spreading, loose;

replum oblong, often constricted, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

subglobose to ellipsoid, uncompressed, 4–6(–9) mm;

valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile;

replum as wide as or wider than fruit;

ovules (8–)10–14(–16) per ovary;

style 1–2.5(–4) mm.

Seeds

flattened.

plump.

2n

= 8, 16.

= 60.

Physaria vitulifera

Physaria arctica

Phenology Flowering Apr–Jun. Flowering May–Aug.
Habitat Rocky hillsides, dry banks, gravel and sand, granitic slopes, soil scree, red shale Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance
Elevation 1600-3000 m (5200-9800 ft) 0-1800 m (0-5900 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia)
[BONAP county map]
Source FNA vol. 7, p. 665. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa
Name authority Rydberg: Bull. Torrey Bot. Club 28: 278. (1901) (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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