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little bladderpod, Moapa bladderpod

Navajo bladderpod

Habit Annuals or, rarely, biennials; with a taproot; densely pubescent, trichomes (simple or stellate, sessile or short-stalked), 4–7-rayed, rays usually furcate, rarely bifurcate, (nearly smooth to finely tuberculate). Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo).
Stems

several from base, decumbent to erect, (several-branched, frequently stout), 1.5–6 dm.

several from base (crowded), erect, not exceeding leaves.

Basal leaves

blade elliptic, (1.5–)3–6.5 cm, margins entire, repand, or shallowly dentate.

usually absent.

Cauline leaves

(proximal shortly petiolate, distal sessile);

blade linear to elliptic or obovate, (0.5–)1–3.5(–4.5) cm, margins entire or repand.

(petiole not differentiated from blade);

blade linear-oblanceolate, 3–8(–13) mm, margins entire.

Racemes

loose.

(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves).

Flowers

sepals oblong, lanceolate, or elliptic, (3–)3.5–6(–7.5) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (yellow to orange), suborbicular or obovate, (5–)6.5–8(–11) mm, (narrowing gradually to broad claw, usually widened at base).

sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate);

petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle).

Fruiting pedicels

(recurved, sigmoid), 5–15 mm.

(ascending to divaricate-ascending, straight), 3.5–6 mm.

Fruits

(sessile or shortly stipitate), orbicular or obovoid, often slightly compressed, (3.5–)4–6 mm;

valves sparsely pubescent, trichomes sessile and stellate, densely pubescent inside, trichomes simple or branched;

ovules 4–12 per ovary;

style 2–4.5 mm.

(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute);

valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not);

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

flattened.

plump or slightly flattened, (strongly mucilaginous).

2n

= 10, 20.

Physaria tenella

Physaria navajoensis

Phenology Flowering Feb–May. Flowering May-early Jun.
Habitat Sandy soils, gravel, clayey loam, loose rocky slopes, washes, desert slopes and plains, lava hills, frequently in or near bushes Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation (0-)600-1900 m ((0-)2000-6200 ft) 2200-2400 m (7200-7900 ft)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 663. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella tenella, Lesquerella gordonii var. sessilis Lesquerella navajoensis
Name authority (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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