Physaria subumbellata |
Physaria hitchcockii |
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parasol bladderpod |
Hitchcock's bladderpod |
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Habit | Perennials; caudex simple or branched, (usually covered with persistent leaf bases, cespitose); densely pubescent, trichomes (closely appressed), rays distinct, usually bifurcate. | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, erect, (unbranched, slender), 0.1–0.6 dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
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Basal leaves | blade rhombic to obovate, 2–4 cm, margins entire. |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
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Cauline leaves | blade linear-oblanceolate, similar to basal. |
similar to basal, smaller. |
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Racemes | dense (distally, subumbellate). |
dense. |
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Flowers | sepals (yellowish), oblong to elliptic, 3.5–7 mm, (median pair usually thickened apically, cucullate); petals lingulate to spatulate, 4–7 mm. |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (divaricate-ascending), 3–5 mm, (densely pubescent). |
(ascending, straight or slightly curved), 2–6 mm. |
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Fruits | (erect), ovate to suborbicular, compressed apically (latiseptate), 3–4 mm; valves pubescent; replum ovate to obovate; ovules 4–6 per ovary; style 2–3 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
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Seeds | plump. |
flattened. |
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2n | = 10. |
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Physaria subumbellata |
Physaria hitchcockii |
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Phenology | Flowering Apr–Jun. | |||||||||
Habitat | Rocky high ridges, gravel and stony areas, juniper covered knolls, rock crevices, clay hillsides, pinyon-juniper areas, calcareous substrates | |||||||||
Elevation | 1600-2700 m (5200-8900 ft) | |||||||||
Distribution |
CO; UT; WY
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NV; UT |
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Discussion | Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 663. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella subumbellata | Lesquerella hitchcockii | ||||||||
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |