Physaria subumbellata |
Physaria didymocarpa |
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parasol bladderpod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; caudex simple or branched, (usually covered with persistent leaf bases, cespitose); densely pubescent, trichomes (closely appressed), rays distinct, usually bifurcate. | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||
Stems | several from base, erect, (unbranched, slender), 0.1–0.6 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | blade rhombic to obovate, 2–4 cm, margins entire. |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | blade linear-oblanceolate, similar to basal. |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | dense (distally, subumbellate). |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals (yellowish), oblong to elliptic, 3.5–7 mm, (median pair usually thickened apically, cucullate); petals lingulate to spatulate, 4–7 mm. |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (divaricate-ascending), 3–5 mm, (densely pubescent). |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | (erect), ovate to suborbicular, compressed apically (latiseptate), 3–4 mm; valves pubescent; replum ovate to obovate; ovules 4–6 per ovary; style 2–3 mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | plump. |
flattened. |
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2n | = 10. |
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Physaria subumbellata |
Physaria didymocarpa |
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Phenology | Flowering Apr–Jun. | |||||||||
Habitat | Rocky high ridges, gravel and stony areas, juniper covered knolls, rock crevices, clay hillsides, pinyon-juniper areas, calcareous substrates | |||||||||
Elevation | 1600-2700 m (5200-8900 ft) | |||||||||
Distribution |
CO; UT; WY
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ID; MT; WA; WY; AB; BC
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 663. | FNA vol. 7, p. 633. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella subumbellata | Vesicaria didymocarpa, Coulterina didymocarpa | ||||||||
Name authority | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||
Web links |