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Fremont County twinpod, Rocky Mountain or Fremont County twinpod

alpine bladderpod, spatula-leaf bladderpod

Habit Perennials; caudex usually simple; (silvery) pubescent throughout, trichome rays furcate. Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate).
Stems

several from base, prostrate to decumbent, 0.3–1 dm.

simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm.

Basal leaves

(rosulate; petiole winged);

blade orbicular to broadly obovate, 1.5–3 cm, margins entire or with broad, obscure toothlike angles each side at apex, (apex obtuse, surfaces densely pubescent, trichomes appressed).

(erect to prostrate, petiole distinct from blade);

blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded).

Cauline leaves

blade broadly spatulate to linear-oblanceolate, 1–1.5 cm, margins entire.

blade spatulate, distinctly different from basal.

Racemes

condensed, (subumbellate to slightly more elongated, few-flowered).

moderately dense, (6–20-flowered).

Flowers

sepals (yellowish, often with some purple), narrowly lanceolate, 5–6 mm;

petals spatulate, 7.3–9.2 mm, (not clawed).

sepals (pale yellow), elliptic, 3.5–5 mm;

petals lingulate, 6–9 mm.

Fruiting pedicels

(divaricate-ascending, straight to slightly curved), 6–10 mm.

(strongly sigmoid), 10–20 mm (2 times longer than fruits).

Fruits

didymous, irregular, suborbicular, deeply bilobed, inflated in age, 10–12 × 12–15 mm, (papery, basal sinus absent or obsolete, apical sinus deep);

valves (retaining seeds after dehiscence), densely pubescent, trichomes spreading, (ovaries and immature fruit downy);

replum narrowly ovate to broadly oblong, not narrowed at middle, as wide as or wider than fruit, apex acute to obtuse;

ovules 4 per ovary;

style 3–7 mm.

lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed);

valves pubescent, trichomes sparse and closely appressed to surface;

ovules 4–8 per ovary;

style 2.5–6 mm (usually ± equal in length to mature fruit).

Seeds

flattened.

plump.

Physaria saximontana

Physaria spatulata

Phenology Flowering May-early Jul.
Habitat Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates
Elevation 900-2900 m (3000-9500 ft)
Distribution
from FNA
MT; WY
[BONAP county map]
from FNA
MT; ND; NE; SD; WY; AB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria saximontana (especially subsp. dentata) is morphologically similar to 22. P. didymocarpa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaf blades: margins entire, apices rounded to angled; styles 3-5 mm.
subsp. saximontana
1. Basal leaf blades: margins dentate (teeth broad), apices angled; styles 4-7 mm.
subsp. dentata
Source FNA vol. 7, p. 661. FNA vol. 7, p. 663.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. saximontana subsp. dentata, P. saximontana subsp. saximontana
Synonyms Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata
Name authority Rollins: Contr. Gray Herb. 214: 13. (1984) (Rydberg) Grady & O’Kane: Novon 17: 190. (2007)
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