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Rollins' twinpod

silver twinpod, west silver bladderpod

Habit Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). Perennials; (diminutive); caudex simple or branched, (buried, with thatch of persistent leaf bases distally); (appearing silvery gray-green to silvery purple), densely pubescent, trichomes usually 5 or 6 (rarely 7)-rayed, rays bifurcate or incompletely so, (relatively short, stout, umbonate, moderately tuberculate to nearly smooth, lower layer smoother).
Stems

several from base, decumbent, (unbranched, slender), 0.5–1 dm.

1–5 from base, prostrate to slightly decumbent, (arising laterally, also erect or ascending from tuft of basal leaves, unbranched, purple-green), 0.08–0.3 dm.

Basal leaves

(strongly rosulate);

blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute).

(petiole slightly winged);

blade oblanceolate, elliptic, or rhombic, (mostly flat, sometimes somewhat folded), 0.6–2.7 cm, (base tapering to petiole), margins entire, (apex rounded to rounded-acute).

Cauline leaves

blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute).

(3–7, shortly petiolate or sessile);

blade elliptic to oblanceolate, 0.3–0.5 cm, margins entire.

Racemes

congested, (elongated moderately in fruit).

crowded, (ca. 3–7 fruits).

Flowers

sepals linear, 5–7 mm;

petals spatulate, 8–10 mm, (apex often somewhat truncate).

sepals (greenish yellow), linear-triangular, 3.7–5 mm, (lateral pair subsaccate);

petals oblanceolate to narrowly obovate, 4.5–9 mm.

Fruiting pedicels

(spreading, straight or somewhat sigmoid), 5–8 mm.

(ascending, straight), 1.8–3.4 mm.

Fruits

(erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit);

replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

(shortly stipitate, purple or greenish purple in age), slightly didymous, ovoid to obpyriform, 3–5 mm (wider than long, base rounded-obtuse, apex rounded, flattened, or slightly emarginate);

valves (inflated, slightly wider than replum), pubescent, trichomes scattered;

replum obovate to orbicular-obdeltate, apex rounded, obtuse, or truncate;

septum complete or medially small-perforate;

ovules 4–6(–8) per ovary;

style 2–3.6 mm.

Seeds

slightly flattened.

relatively plump, (ovate to suborbicular, usually rounded on one side, ± flat or concave on the other, not mucilaginous when wetted).

2n

= 8.

Physaria rollinsii

Physaria scrotiformis

Phenology Flowering May–Jun. Flowering Jun-early Jul.
Habitat Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders Tundra areas with islands of Engelmann spruce on Leadville limestone, amidst limestone cobbles and gravel
Elevation 2300-2500(-3900) m (7500-8200(-12800) ft) 3500-3700 m (11500-12100 ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
CO
[BONAP county map]
Discussion

Physaria scrotiformis is known only from La Plata County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 661. FNA vol. 7, p. 662.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Name authority G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) O’Kane: Novon 17: 376, fig. 1. (2007)
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