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Rollins' twinpod

Hitchcock's bladderpod

Habit Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate).
Stems

several from base, decumbent, (unbranched, slender), 0.5–1 dm.

few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm.

Basal leaves

(strongly rosulate);

blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute).

(petiole and blade differentiated or not);

blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire.

Cauline leaves

blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute).

similar to basal, smaller.

Racemes

congested, (elongated moderately in fruit).

dense.

Flowers

sepals linear, 5–7 mm;

petals spatulate, 8–10 mm, (apex often somewhat truncate).

sepals narrowly lanceolate to lanceolate, 2.8–6 mm;

petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade).

Fruiting pedicels

(spreading, straight or somewhat sigmoid), 5–8 mm.

(ascending, straight or slightly curved), 2–6 mm.

Fruits

(erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep);

valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit);

replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse;

ovules 4 per ovary;

style 5–7 mm.

(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute);

valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.7–6 mm.

Seeds

slightly flattened.

flattened.

2n

= 8.

Physaria rollinsii

Physaria hitchcockii

Phenology Flowering May–Jun.
Habitat Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders
Elevation 2300-2500(-3900) m (7500-8200(-12800) ft)
Distribution
from FNA
CO
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation).
subsp. rubicundula
1. Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah
→ 2
2. Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada.
subsp. hitchcockii
2. Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada.
subsp. confluens
Source FNA vol. 7, p. 661. FNA vol. 7, p. 643.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hitchcockii subsp. confluens, P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
Synonyms Lesquerella hitchcockii
Name authority G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002)
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