Physaria rollinsii |
Physaria hitchcockii |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
Rollins' twinpod |
Hitchcock's bladderpod |
|||||||||
Habit | Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, decumbent, (unbranched, slender), 0.5–1 dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
||||||||
Basal leaves | (strongly rosulate); blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute). |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
||||||||
Cauline leaves | blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute). |
similar to basal, smaller. |
||||||||
Racemes | congested, (elongated moderately in fruit). |
dense. |
||||||||
Flowers | sepals linear, 5–7 mm; petals spatulate, 8–10 mm, (apex often somewhat truncate). |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
||||||||
Fruiting pedicels | (spreading, straight or somewhat sigmoid), 5–8 mm. |
(ascending, straight or slightly curved), 2–6 mm. |
||||||||
Fruits | (erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit); replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
||||||||
Seeds | slightly flattened. |
flattened. |
||||||||
2n | = 8. |
|||||||||
Physaria rollinsii |
Physaria hitchcockii |
|||||||||
Phenology | Flowering May–Jun. | |||||||||
Habitat | Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders | |||||||||
Elevation | 2300-2500(-3900) m (7500-8200(-12800) ft) | |||||||||
Distribution |
CO
|
NV; UT |
||||||||
Discussion | Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|||||||||
Key |
|
|||||||||
Source | FNA vol. 7, p. 661. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella hitchcockii | |||||||||
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |