Physaria rollinsii |
Physaria didymocarpa |
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Rollins' twinpod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||
Stems | several from base, decumbent, (unbranched, slender), 0.5–1 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | (strongly rosulate); blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute). |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute). |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | congested, (elongated moderately in fruit). |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals linear, 5–7 mm; petals spatulate, 8–10 mm, (apex often somewhat truncate). |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (spreading, straight or somewhat sigmoid), 5–8 mm. |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | (erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit); replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | slightly flattened. |
flattened. |
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2n | = 8. |
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Physaria rollinsii |
Physaria didymocarpa |
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Phenology | Flowering May–Jun. | |||||||||
Habitat | Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders | |||||||||
Elevation | 2300-2500(-3900) m (7500-8200(-12800) ft) | |||||||||
Distribution |
CO
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ID; MT; WA; WY; AB; BC
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Discussion | Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 661. | FNA vol. 7, p. 633. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Vesicaria didymocarpa, Coulterina didymocarpa | |||||||||
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||
Web links |