Physaria rollinsii |
Physaria chambersii |
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Rollins' twinpod |
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
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Habit | Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). |
Stems | several from base, decumbent, (unbranched, slender), 0.5–1 dm. |
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
Basal leaves | (strongly rosulate); blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute). |
(petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
Cauline leaves | blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute). |
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
Racemes | congested, (elongated moderately in fruit). |
congested. |
Flowers | sepals linear, 5–7 mm; petals spatulate, 8–10 mm, (apex often somewhat truncate). |
sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
Fruiting pedicels | (spreading, straight or somewhat sigmoid), 5–8 mm. |
(divaricate, slightly sigmoid), 8–15 mm. |
Fruits | (erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit); replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
Seeds | slightly flattened. |
flattened. |
2n | = 8. |
= 8, 10, 16, 24. |
Physaria rollinsii |
Physaria chambersii |
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Phenology | Flowering May–Jun. | Flowering Apr–Jul. |
Habitat | Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas |
Elevation | 2300-2500(-3900) m (7500-8200(-12800) ft) | 1500-3200 m (4900-10500 ft) |
Distribution |
CO
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AZ; CA; ID; NV; OR; UT
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Discussion | Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 661. | FNA vol. 7, p. 631. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. chambersii var. canaani, P. chambersii var. sobolifera | |
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) |
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