Physaria rollinsii |
Physaria arctica |
|
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Rollins' twinpod |
arctic bladderpod |
|
Habit | Perennials; (compact); caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes 6–8-rayed, rays furcate near base, fused at base, (umbonate, strongly tuberculate throughout). | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). |
Stems | several from base, decumbent, (unbranched, slender), 0.5–1 dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
Basal leaves | (strongly rosulate); blade usually oblanceolate or broader, sometimes triangular, 2–3.5 cm (width 5–10 mm), margins entire or with 1 or 2 broad teeth, (apex acute). |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
Cauline leaves | blade oblanceolate, 1–1.5 cm (width 2–4 mm), margins entire, (apex acute). |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
Racemes | congested, (elongated moderately in fruit). |
loose. |
Flowers | sepals linear, 5–7 mm; petals spatulate, 8–10 mm, (apex often somewhat truncate). |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
Fruiting pedicels | (spreading, straight or somewhat sigmoid), 5–8 mm. |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
Fruits | (erect), didymous, suborbicular, inflated, 2–5(–8) × 4–8(–10)mm, (coriaceous, base slightly cordate or nearly obtuse, sinus obsolete or absent, apical sinus broad and deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed, (silvery on ovaries and immature fruit); replum obovate to oblong, as wide as or wider than fruit, rarely somewhat constricted basally, apex obtuse; ovules 4 per ovary; style 5–7 mm. |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
Seeds | slightly flattened. |
plump. |
2n | = 8. |
= 60. |
Physaria rollinsii |
Physaria arctica |
|
Phenology | Flowering May–Jun. | Flowering May–Aug. |
Habitat | Sagebrush, granitic talus, open knolls, limestone chiprock, steep slopes, clay banks, near granite boulders | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance |
Elevation | 2300-2500(-3900) m (7500-8200(-12800) ft) | 0-1800 m (0-5900 ft) |
Distribution |
CO
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AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
Source | FNA vol. 7, p. 661. | FNA vol. 7, p. 626. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa | |
Name authority | G. A. Mulligan: Canad. J. Bot. 44: 1663, fig. 2, plate 1, fig. 4. (1966) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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