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alpine bladderpod, reed's twinpod, Rollins' bladderpod

straight bladderpod

Habit Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). Perennials; (loose, spreading); caudex simple or branched; densely pubescent, trichomes (subsessile), 4–6-rayed, rays furcate or bifurcate, (moderately tuberculate over arms, less so or smooth over center).
Stems

few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm.

few to several from base, ascending or prostrate, (arising laterally, also from within basal leaves, usually unbranched, rarely branched), 0.5–3(–6) dm.

Basal leaves

(erect);

blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire.

blade narrowly oblanceolate or broadly elliptic, 1–7(–12) cm, margins entire or shallowly toothed, sometimes repand, (inner blades usually flattened in age, surfaces often gray-green, scabrous).

Cauline leaves

blade linear, similar to basal.

(usually secund);

blade spatulate or obovate, 1–2.5(–4.5) cm, margins entire or shallowly toothed, (flat or involute).

Racemes

dense, (often subumbellate, not or barely exceeding basal leaves).

somewhat crowded (to moderately elongated in fruit, exceeding basal leaves).

Flowers

sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals lingulate, 6–9 mm.

sepals broadly elliptic or oblong, 4–7.5(–9) mm, (median pair thickened apically, cucullate);

petals cuneate or obovate, 7–10(–16) mm, (tapering gradually to broad claw).

Fruiting pedicels

(ascending, curved), 3–5.5 mm.

(often divaricate-spreading and straight, or horizontal and loosely sigmoid, sometimes slightly recurved), 5–15 mm.

Fruits

lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm;

valves pubescent, trichomes closely appressed;

ovules 8–12 per ovary;

style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma).

subglobose to ovoid or ellipsoid, sometimes compressed, (4–)5–7(–9) mm;

valves sparsely pubescent, trichomes appressed or erect, sometimes sparsely pubescent inside;

ovules (8–)12–16(–20) per ovary;

style 2–7 mm.

Seeds

plump, (oblong).

somewhat flattened.

2n

= 10 + 2, 18, 20, ± 40.

Physaria reediana

Physaria rectipes

Phenology Flowering May–Jul. Flowering May–Jul.
Habitat Open areas of grasslands on calcareous soils Sandy soils, limey knolls, rocky hills, clay hillsides, dry ridges, weathered rocks, gravelly outwashes, stony slopes, pinyon-juniper woodlands
Elevation 1200-1900 m (3900-6200 ft) 1500-2600 m (4900-8500 ft)
Distribution
from FNA
CO; NE; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

As here circumscribed, Physaria rectipes remains heterogeneous and may represent more than one taxon, even after the recent removal of P. pulvinata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 660. FNA vol. 7, p. 660.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis Lesquerella rectipes
Name authority O’Kane & Al-Shehbaz: Novon 12: 327. (2002) (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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