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alpine bladderpod, reed's twinpod, Rollins' bladderpod

Graham's twinpod

Habit Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate.
Stems

few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm.

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

Basal leaves

(erect);

blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire.

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

Cauline leaves

blade linear, similar to basal.

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

Racemes

dense, (often subumbellate, not or barely exceeding basal leaves).

loose, (elongated).

Flowers

sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate);

petals lingulate, 6–9 mm.

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

Fruiting pedicels

(ascending, curved), 3–5.5 mm.

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

Fruits

lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm;

valves pubescent, trichomes closely appressed;

ovules 8–12 per ovary;

style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma).

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

Seeds

plump, (oblong).

plump, (suborbicular).

Physaria reediana

Physaria grahamii

Phenology Flowering May–Jul. Flowering May–Jun.
Habitat Open areas of grasslands on calcareous soils Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay
Elevation 1200-1900 m (3900-6200 ft) 2100-2900 m (6900-9500 ft)
Distribution
from FNA
CO; NE; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 660. FNA vol. 7, p. 642.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda
Name authority O’Kane & Al-Shehbaz: Novon 12: 327. (2002) C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937)
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