Physaria reediana |
Physaria grahamii |
|
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alpine bladderpod, reed's twinpod, Rollins' bladderpod |
Graham's twinpod |
|
Habit | Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | (erect); blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | blade linear, similar to basal. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | dense, (often subumbellate, not or barely exceeding basal leaves). |
loose, (elongated). |
Flowers | sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals lingulate, 6–9 mm. |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (ascending, curved), 3–5.5 mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm; valves pubescent, trichomes closely appressed; ovules 8–12 per ovary; style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma). |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | plump, (oblong). |
plump, (suborbicular). |
Physaria reediana |
Physaria grahamii |
|
Phenology | Flowering May–Jul. | Flowering May–Jun. |
Habitat | Open areas of grasslands on calcareous soils | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 1200-1900 m (3900-6200 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
CO; NE; WY
|
UT |
Discussion | Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 660. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |