Physaria reediana |
Physaria bellii |
|
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alpine bladderpod, reed's twinpod, Rollins' bladderpod |
Bell's or Front Range twinpod, Bell's twinpod, Front Range twinpod |
|
Habit | Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex simple, (relatively large); densely (silvery) pubescent, trichomes (sessile, appressed), rays furcate, fused at base. |
Stems | few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm. |
simple from base, decumbent to nearly prostrate, 0.5–1.3 dm. |
Basal leaves | (erect); blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire. |
(strongly rosulate; shortly petiolate); blade broadly obovate, 1.5–7.5 (width 7.5–26 mm, base gradually tapering to petiole), margins shallowly dentate, (apex obtuse). |
Cauline leaves | blade linear, similar to basal. |
blade oblanceolate to broadly obovate, 1–2.5 cm, margins entire. |
Racemes | dense, (often subumbellate, not or barely exceeding basal leaves). |
dense. |
Flowers | sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals lingulate, 6–9 mm. |
sepals (pale yellow or yellow-green), narrowly lanceolate to narrowly deltate, 4–8 mm; petals yellow, broadly spatulate to obovate, 9–13 mm, (not clawed). |
Fruiting pedicels | (ascending, curved), 3–5.5 mm. |
(divaricate-ascending to widely spreading, slightly sigmoid to curved), 7–12 mm. |
Fruits | lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm; valves pubescent, trichomes closely appressed; ovules 8–12 per ovary; style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma). |
didymous, slightly flattened (contrary to replum) to uncompressed, 4–9 × 2–8 mm, (strongly coriaceous, apical and basal sinuses narrow, deep); valves (retaining seeds after dehiscence), pubescent, trichomes appressed; replum narrowly oblanceolate to narrowly linear-oblong, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style more than 3 mm. |
Seeds | plump, (oblong). |
compressed. |
2n | = 8. |
|
Physaria reediana |
Physaria bellii |
|
Phenology | Flowering May–Jul. | Flowering Mar–Jun(-Jul). |
Habitat | Open areas of grasslands on calcareous soils | Dark shale, road cuts, ridge crests, washes |
Elevation | 1200-1900 m (3900-6200 ft) | 1500-1800 m (4900-5900 ft) |
Distribution |
CO; NE; WY
|
CO
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Discussion | Of conservation concern. Physaria bellii is often found in shale and limestone soils of the Fountain/Ingleside, Lykins, Niobrara, and Pierre formations. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 660. | FNA vol. 7, p. 628. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis | |
Name authority | O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | G. A. Mulligan: Canad. J. Bot. 44: 1662, fig. 1, plate 1, fig. 3. (1966) |
Web links |