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rose bladderpod

Navajo bladderpod

Habit Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo).
Stems

simple from base, erect, (unbranched, sparsely leaved), to 7 dm.

several from base (crowded), erect, not exceeding leaves.

Basal leaves

blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid.

usually absent.

Cauline leaves

(proximal often narrowed to short petiole, distal sessile);

blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire.

(petiole not differentiated from blade);

blade linear-oblanceolate, 3–8(–13) mm, margins entire.

Racemes

dense or slightly elongated.

(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves).

Flowers

sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate);

petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade).

sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate);

petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle).

Fruiting pedicels

(spreading or recurved, loosely sigmoid), 5–25 mm.

(ascending to divaricate-ascending, straight), 3.5–6 mm.

Fruits

(pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8(–12) per ovary;

style 1–3(–4) mm.

(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute);

valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not);

ovules 4–8 per ovary;

style 1.8–3 mm.

Seeds

flattened.

plump or slightly flattened, (strongly mucilaginous).

2n

= 18, 36.

Physaria purpurea

Physaria navajoensis

Phenology Flowering Mar–Oct. Flowering May-early Jun.
Habitat Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone
Elevation 400-2400 m (1300-7900 ft) 2200-2400 m (7200-7900 ft)
Distribution
from FNA
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Of conservation concern.

Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 659. FNA vol. 7, p. 651.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa Lesquerella navajoensis
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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