Physaria purpurea |
Physaria integrifolia |
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rose bladderpod |
Snake River or creeping twinpod, Snake River twinpod |
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Habit | Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). | Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout). |
Stems | simple from base, erect, (unbranched, sparsely leaved), to 7 dm. |
several from base, ± erect, exceeding basal rosette by ± 0.5 dm. |
Basal leaves | blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid. |
(forming a strong rosette; long-petiolate); blade oblanceolate to ovate or orbicular, (1.5–)2–4(–8) cm, (base usually abruptly tapering to petiole), margins entire. |
Cauline leaves | (proximal often narrowed to short petiole, distal sessile); blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire. |
blade oblanceolate, 1–2 cm, margin entire, (apex acute). |
Racemes | dense or slightly elongated. |
congested, (greatly exceeding leaves). |
Flowers | sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate); petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade). |
sepals often keeled, 6–8 mm; petals spatulate, 8–10 mm. |
Fruiting pedicels | (spreading or recurved, loosely sigmoid), 5–25 mm. |
(spreading, straight or slightly curved), 7–11 mm. |
Fruits | (pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 1–3(–4) mm. |
didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed; replum linear to oblong, as wide as or wider than fruit; ovules 8 per ovary; style 7–9 mm. |
Seeds | flattened. |
flattened. |
2n | = 18, 36. |
= 16. |
Physaria purpurea |
Physaria integrifolia |
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Phenology | Flowering Mar–Oct. | Flowering (May-)Jun–Jul(-Aug). |
Habitat | Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps | Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale |
Elevation | 400-2400 m (1300-7900 ft) | 1900-2700 m (6200-8900 ft) |
Distribution |
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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ID; MT; WY |
Discussion | Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 659. | FNA vol. 7, p. 644. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa | P. didymocarpa var. integrifolia, P. integrifolia var. monticola |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | (Rollins) Lichvar: Madroño 31: 203. (1984) |
Web links |