Physaria purpurea |
Physaria arizonica |
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rose bladderpod |
Arizona bladderpod |
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Habit | Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). |
Stems | simple from base, erect, (unbranched, sparsely leaved), to 7 dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
Basal leaves | blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid. |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
Cauline leaves | (proximal often narrowed to short petiole, distal sessile); blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire. |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
Racemes | dense or slightly elongated. |
dense, often subcorymbiform. |
Flowers | sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate); petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade). |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
Fruiting pedicels | (spreading or recurved, loosely sigmoid), 5–25 mm. |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
Fruits | (pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 1–3(–4) mm. |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
Seeds | flattened. |
flattened. |
2n | = 18, 36. |
= 10. |
Physaria purpurea |
Physaria arizonica |
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Phenology | Flowering Mar–Oct. | Flowering Apr–Jun. |
Habitat | Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine |
Elevation | 400-2400 m (1300-7900 ft) | 1000-2200 m (3300-7200 ft) |
Distribution |
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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AZ; UT
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Discussion | The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 659. | FNA vol. 7, p. 627. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
Web links |