Physaria purpurea |
Physaria arctica |
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rose bladderpod |
arctic bladderpod |
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Habit | Perennials; caudex simple, (usually woody); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, (smooth or tuberculate). | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). |
Stems | simple from base, erect, (unbranched, sparsely leaved), to 7 dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
Basal leaves | blade elliptic or obovate to oblong, 4–15 cm, margins entire, dentate, or lyrate-pinnatifid. |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
Cauline leaves | (proximal often narrowed to short petiole, distal sessile); blade broadly elliptic to obovate or rhombic, 0.5–3(–5) cm, margins entire. |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
Racemes | dense or slightly elongated. |
loose. |
Flowers | sepals elliptic to ovate, 3.5–6(–7) mm, (median pair usually thickened apically, cucullate); petals (white, often purple-veined, fading purplish), suborbicular to obovate, obdeltate, or cuneate, 4.5–10(–12) mm, (often narrowed to broad claw, apex emarginate, less frequently claw undifferentiated from blade). |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
Fruiting pedicels | (spreading or recurved, loosely sigmoid), 5–25 mm. |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
Fruits | (pendent or horizontal, sessile or substipitate), subglobose to broadly ellipsoid, not or slightly inflated, (4–)5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 1–3(–4) mm. |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
Seeds | flattened. |
plump. |
2n | = 18, 36. |
= 60. |
Physaria purpurea |
Physaria arctica |
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Phenology | Flowering Mar–Oct. | Flowering May–Aug. |
Habitat | Rocky draws, canyons, stony hills, ridges, rock crevices on limestone ledges, lava cliffs, sand and gravel of dry stream beds, rocky slopes, talus, shade of bushes or cactus clumps | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance |
Elevation | 400-2400 m (1300-7900 ft) | 0-1800 m (0-5900 ft) |
Distribution |
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
Source | FNA vol. 7, p. 659. | FNA vol. 7, p. 626. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria purpurea, Lesquerella purpurea, Lesquerella purpurea subsp. foliosa, Lesquerella purpurea var. foliosa, P. purpurea var. foliosa | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa |
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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