Physaria pruinosa |
Physaria ludoviciana |
|
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frosty bladderpod, Pagosa bladderpod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; caudex simple or branched, (covered with persistent leaf bases); densely pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). |
Stems | simple or several from base, decumbent or erect, (unbranched), to 2 dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
Basal leaves | (petiole sharply differentiated from blade, slender); blade suborbicular or obovate to rhombic, 4–8 cm, (base abruptly narrowed to petiole), margins entire, sinuate, or shallowly dentate, (abaxial surface densely pubescent, adaxial lightly pubescent). |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
Cauline leaves | (proximal petiolate, distal sessile); blade obovate to rhombic, 0.8–2.3 cm, margins entire or shallowly toothed. |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
Racemes | dense, (somewhat elongated in fruit). |
compact, (elongated and loose in fruit, densely-flowered). |
Flowers | sepals elliptic or oblong, ca. 6 mm, (lateral pair not saccate or subsaccate, cucullate, median pair thickened apically, cucullate); petals spatulate, ca. 9 mm, (claw expanded at base). |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
Fruiting pedicels | (horizontal to ascending, sigmoid or slightly curved), 8–11 mm, (stout). |
(usually recurved), (5–)10–20(–25) mm. |
Fruits | (sessile or substipitate, often becoming copper-red in age), subglobose or ellipsoid, inflated, 6–9 mm, (firm, glossy); valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 3.5–7 mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
Seeds | somewhat flattened. |
slightly flattened. |
2n | = 10, 20, 30. |
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Physaria pruinosa |
Physaria ludoviciana |
|
Phenology | Flowering May–Jun(-Aug). | Flowering Apr–Jul(-Aug). |
Habitat | Mancos slate or shale, meadows, gentle slopes, edges of ponderosa pine stands | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands |
Elevation | 2100-2600 m (6900-8500 ft) | 0-1900 m (0-6200 ft) |
Distribution |
CO; NM |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Of conservation concern. The one New Mexico population is near the border with Colorado, in Rio Arriba County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 658. | FNA vol. 7, p. 649. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Lesquerella pruinosa | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana |
Name authority | (Greene) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |