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frosty bladderpod, Pagosa bladderpod

foothill bladderpod, silver bladderpod

Habit Perennials; caudex simple or branched, (covered with persistent leaf bases); densely pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout).
Stems

simple or several from base, decumbent or erect, (unbranched), to 2 dm.

few from base, erect with outer usually decumbent, 1–3.5(–5) dm.

Basal leaves

(petiole sharply differentiated from blade, slender);

blade suborbicular or obovate to rhombic, 4–8 cm, (base abruptly narrowed to petiole), margins entire, sinuate, or shallowly dentate, (abaxial surface densely pubescent, adaxial lightly pubescent).

(erect);

blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes).

Cauline leaves

(proximal petiolate, distal sessile);

blade obovate to rhombic, 0.8–2.3 cm, margins entire or shallowly toothed.

blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute.

Racemes

dense, (somewhat elongated in fruit).

compact, (elongated and loose in fruit, densely-flowered).

Flowers

sepals elliptic or oblong, ca. 6 mm, (lateral pair not saccate or subsaccate, cucullate, median pair thickened apically, cucullate);

petals spatulate, ca. 9 mm, (claw expanded at base).

sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate);

petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw).

Fruiting pedicels

(horizontal to ascending, sigmoid or slightly curved), 8–11 mm, (stout).

(usually recurved), (5–)10–20(–25) mm.

Fruits

(sessile or substipitate, often becoming copper-red in age), subglobose or ellipsoid, inflated, 6–9 mm, (firm, glossy);

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8(–12) per ovary;

style 3.5–7 mm.

subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm;

valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile;

ovules (4–)8–12(–16) per ovary;

style 3–4.5(–6.5) mm.

Seeds

somewhat flattened.

slightly flattened.

2n

= 10, 20, 30.

Physaria pruinosa

Physaria ludoviciana

Phenology Flowering May–Jun(-Aug). Flowering Apr–Jul(-Aug).
Habitat Mancos slate or shale, meadows, gentle slopes, edges of ponderosa pine stands Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands
Elevation 2100-2600 m (6900-8500 ft) 0-1900 m (0-6200 ft)
Distribution
from FNA
CO; NM
[BONAP county map]
from FNA
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

The one New Mexico population is near the border with Colorado, in Rio Arriba County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 658. FNA vol. 7, p. 649.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella pruinosa Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana
Name authority (Greene) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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