Physaria pinetorum |
Physaria grahamii |
|
---|---|---|
White Mountain bladderpod |
Graham's twinpod |
|
Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | (petiole tapering to blade); blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire. |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | (not or loosely overlapping, petiolate or distal sessile); blade spatulate to oblanceolate, 1–4 cm, margins entire. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | crowded, elongated. |
loose, (elongated). |
Flowers | sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate); petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (ascending, curved or sigmoid), 6–12(–20) mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | (substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–24 per ovary; style (2–)4–7 mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | flattened. |
plump, (suborbicular). |
2n | = 10. |
|
Physaria pinetorum |
Physaria grahamii |
|
Phenology | Flowering Apr–Jul. | Flowering May–Jun. |
Habitat | Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 1400-2900 (-3400) m (4600-9500 (-11200) ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
AZ; NM
|
UT |
Discussion | Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 658. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Lesquerella pinetorum | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
Web links |