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thick-leaf bladderpod

Mcvaugh's bladderpod

Habit Perennials; caudex usually simple, rarely branched, (loosely mounded, rosette-like growth); densely (silvery or gray) pubescent, trichomes (sessile), 5-rayed, rays bifurcate, slightly fused near base of main rays, (tuberculate throughout, less over umbo). Perennials; caudex simple or branched, (sometimes enlarged); densely pubescent, trichomes (sessile), several-rayed, rays fused (webbed) most of their length, (umbonate, peltate, tuberculate throughout).
Stems

several from base, decumbent to prostrate, (well-exserted beyond basal leaves), 0.2–0.5 dm.

few to several from base, erect or outer ones decumbent, 0.5–4 dm.

Basal leaves

(petiole differentiated from blade);

blade (slightly cupped, leathery, nearly 1 mm thick), oblanceolate to orbicular, 1.2–2 cm, margins entire, (apex acute).

(long-petiolate);

blade elliptic to obovate or rhombic, 2–6(–9) cm, margins entire.

Cauline leaves

blade spatulate, similar to basal.

(sessile or shortly petiolate);

blade oblanceolate to spatulate, 1–3 cm, (proximal broader), margins entire.

Racemes

dense, (subumbellate).

dense, (relatively short).

Flowers

sepals (pale yellow), elliptic to oblong, 3.5–4.0 mm, (median pair somewhat thickened apically, cucullate);

petals lingulate, 5–6 mm.

sepals elliptic or narrowly oblong, 4–5.4 mm, (tapered to apex);

petals (white, base and claw yellow, conspicuously purple-veined), usually broadly obovate or rhombic, 6–10 mm, (± equal to blade, tapering to slender claw).

Fruiting pedicels

(ascending, curved), 5–7 mm.

(erect to spreading, ascending, or (proximal) horizontal, straight to slightly curved, sometimes loosely sigmoid), 6–12(–20) mm.

Fruits

globose or ellipsoid, slightly inflated (with slight apical constriction), 3–6 mm;

valves pubescent, trichomes closely appressed;

ovules 8 per ovary;

style 1–3 mm (shorter than mature fruit).

(sessile or substipitate, often reddish magenta), usually ovoid to subglobose, inflated, 4–6(–7) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

septum perforate;

ovules 8–12 per ovary;

style 1.5–4 mm.

Seeds

plump, (oblong).

somewhat flattened.

2n

= 12.

Physaria pachyphylla

Physaria mcvaughiana

Phenology Flowering Jun–Jul. Flowering mid Mar–Apr(-Aug).
Habitat Barren areas of mixed white, pink, or reddish limestone and diatomaceous earth Stream bed gravels, rocky limestone slopes and hills, canyon bottoms and slopes, limestone rubble
Elevation 1300-1600 m (4300-5200 ft) 1200-1600 m (3900-5200 ft)
Distribution
from FNA
MT
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Of conservation concern.

Physaria pachyphylla is known from the Pryor Mountain Desert near the Wyoming state line.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 656. FNA vol. 7, p. 650.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella mcvaughiana
Name authority O’Kane & Grady: Novon 17: 187, fig. 4. (2007) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002)
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