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thick-leaf bladderpod

limestone glade or Missouri bladderpod

Habit Perennials; caudex usually simple, rarely branched, (loosely mounded, rosette-like growth); densely (silvery or gray) pubescent, trichomes (sessile), 5-rayed, rays bifurcate, slightly fused near base of main rays, (tuberculate throughout, less over umbo). Annuals; with a fine taproot; densely pubescent, trichomes (sessile), usually 4-rayed, rays forked, rarely simple or tripartite, (finely tuberculate).
Stems

several from base, decumbent to prostrate, (well-exserted beyond basal leaves), 0.2–0.5 dm.

several from base, (slender), erect or outer decumbent, (usually branched, branches filiform, bud clusters of growing plants drooping), to 2.5 dm.

Basal leaves

(petiole differentiated from blade);

blade (slightly cupped, leathery, nearly 1 mm thick), oblanceolate to orbicular, 1.2–2 cm, margins entire, (apex acute).

blade orbicular to broadly spatulate, 1–2.4 cm, margins entire or sinuate.

Cauline leaves

blade spatulate, similar to basal.

(proximal often petiolate, distal sessile);

similar to basal, blade spatulate to oblanceolate or (distal) linear, (base cuneate), margins entire or sinuate.

Racemes

dense, (subumbellate).

loose.

Flowers

sepals (pale yellow), elliptic to oblong, 3.5–4.0 mm, (median pair somewhat thickened apically, cucullate);

petals lingulate, 5–6 mm.

sepals oblong or elliptic, 2.5–4.6 mm, (median pair slightly thickened apically);

petals (pale yellow), spatulate to obovate, 5–9 mm, (apex ± emarginate).

Fruiting pedicels

(ascending, curved), 5–7 mm.

(usually divaricate-ascending, straight or slightly curved), 7–11 mm.

Fruits

globose or ellipsoid, slightly inflated (with slight apical constriction), 3–6 mm;

valves pubescent, trichomes closely appressed;

ovules 8 per ovary;

style 1–3 mm (shorter than mature fruit).

(sessile or shortly stipitate), globose, not inflated, 3–4 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4 per ovary;

style 3–5 mm.

Seeds

plump, (oblong).

flattened.

2n

= 14.

Physaria pachyphylla

Physaria filiformis

Phenology Flowering Jun–Jul. Flowering Apr–May.
Habitat Barren areas of mixed white, pink, or reddish limestone and diatomaceous earth Limestone, dolomite, and shale, sparsely vegetated or barren areas, cedar glades, old pastures, along roadsides
Elevation 1300-1600 m (4300-5200 ft) 200-300 m (700-1000 ft)
Distribution
from FNA
MT
[BONAP county map]
from FNA
AL; AR; MO
[BONAP county map]
Discussion

Of conservation concern.

Physaria pachyphylla is known from the Pryor Mountain Desert near the Wyoming state line.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 656. FNA vol. 7, p. 637.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella filiformis
Name authority O’Kane & Grady: Novon 17: 187, fig. 4. (2007) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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