Physaria ovalifolia |
Physaria newberryi |
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roundleaf bladderpod |
Newberry twinpod, Newberry's twinpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | ||||||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
(ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
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Seeds | flattened. |
slightly flattened, (ovate). |
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Physaria ovalifolia |
Physaria newberryi |
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Distribution |
CO; KS; NE; NM; OK; TX
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AZ; NM; NV; UT
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 652. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
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Subordinate taxa | ||||||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | P. didymocarpa var. newberryi | ||||||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | ||||||||
Web links |