Physaria ovalifolia |
Physaria navajoensis |
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roundleaf bladderpod |
Navajo bladderpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Perennials; caudex branched, (woody, pulvinate-cespitose, forming hard, hemispherical mats, basal parts covered with persistent leaf bases); densely (silvery gray) pubescent, trichomes mostly 5-rayed, rays bifurcate, slightly fused at base, (umbonate, strongly tuberculate except nearly smooth over umbo). | ||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
several from base (crowded), erect, not exceeding leaves. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
usually absent. |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
(petiole not differentiated from blade); blade linear-oblanceolate, 3–8(–13) mm, margins entire. |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
(secund), dense, corymbose, (few-flowered, not or barely exceeding leaves). |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals (yellow-green), linear to narrowly triangular, 3.7–4.8 mm, (lateral pair subsaccate); petals (deep yellow, slightly orange in center), spatulate, 5.2–6.5 mm, (claw joined at right angle). |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(ascending to divaricate-ascending, straight), 3.5–6 mm. |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
(becoming reddish or copper-colored in age), ovate, often slightly compressed (at margins apically), 3–5 mm, (apex acute); valves (not retaining seeds after dehiscence), glabrous; (septum perforate or not); ovules 4–8 per ovary; style 1.8–3 mm. |
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Seeds | flattened. |
plump or slightly flattened, (strongly mucilaginous). |
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Physaria ovalifolia |
Physaria navajoensis |
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Phenology | Flowering May-early Jun. | |||||
Habitat | Pinyon-juniper communities on nearly barren outcrops of Todilto Limestone | |||||
Elevation | 2200-2400 m (7200-7900 ft) | |||||
Distribution |
CO; KS; NE; NM; OK; TX
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AZ; NM |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria navajoensis is morphologically similar to 85. P. tumulosa of southern Utah, differing subtly. Physaria navajoensis has petals slightly orange at the junction of blade and claw, a sharp bend at that junction giving the flower a flat-topped appearance, and strongly mucilaginous seeds. Physaria tumulosa has pure yellow petals that gently flex at the junction of blade and claw, and seeds that are not mucilaginous. Molecular data (pers. obs.) show that these two species are not directly related. A population of plants on Deer Spring Point, Kane County, Utah, appears to be this species, but molecular data indicate that it is probably a hybrid between P. tumulosa and, most likely, P. intermedia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 651. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | Lesquerella navajoensis | ||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | (O’Kane) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||
Web links |