Physaria ovalifolia |
Physaria ludoviciana |
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roundleaf bladderpod |
foothill bladderpod, silver bladderpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | ||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
(erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
compact, (elongated and loose in fruit, densely-flowered). |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(usually recurved), (5–)10–20(–25) mm. |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
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Seeds | flattened. |
slightly flattened. |
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2n | = 10, 20, 30. |
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Physaria ovalifolia |
Physaria ludoviciana |
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Phenology | Flowering Apr–Jul(-Aug). | |||||
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | |||||
Elevation | 0-1900 m (0-6200 ft) | |||||
Distribution |
CO; KS; NE; NM; OK; TX
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AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 649. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | ||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||
Web links |