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roundleaf bladderpod

Kane County twinpod

Habit Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). Perennials; caudex simple, (with deep roots, thickened); densely (silvery) pubescent throughout (densely covering leaves with several appressed layers), less dense on stems, trichomes (stellate-scalelike), rays fused (webbed) in proximal 1/2 or to tips, (umbonate, nearly smooth to moderately tuberculate).
Stems

few to several from base, erect or outer decumbent, 0.5–2.5 dm.

simple from base, erect or outer ones slightly decumbent toward base, (from below or in basal leaves, unbranched), (0.5–)0.8–1.6(–2) dm.

Basal leaves

blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate.

(erect, petiole long, slender);

blade spatulate to broadly oblanceolate, (3–)5–7(–12) cm, (base gradually tapering to petiole), margins entire, (apex rounded or obtuse).

Cauline leaves

(proximal shortly petiolate, distal usually sessile);

blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire.

blade oblanceolate, similar to basal, (base cuneate), margins entire.

Racemes

compact, (± subumbellate to densely corymbiform, elongated or not).

dense.

Flowers

sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically);

petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated).

sepals (erect), linear to linear-oblong, somewhat boat-shaped, 7–10 mm;

petals (erect at anthesis), lingulate, 11–15 mm, (claw undifferentiated from blade).

Fruiting pedicels

(usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout).

(divaricate-ascending, straight or slightly curved), 10–15 mm.

Fruits

(sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 4–8(–9) mm.

(purplish in age), strongly didymous, semiorbicular, highly inflated, 10–18 × 14–19 mm, (papery), basal sinus usually shallow, rarely absent, apical sinus deep, narrowly V-shaped;

valves retaining seeds after dehiscence, sides flat, back rounded, margins keeled, base and apex obtuse;

replum narrowly oblong to linear, as wide as or wider than fruit, base slightly narrowed, apex obtusely rounded;

ovules 4 per ovary;

style 3–5 mm, (slender).

Seeds

flattened.

slightly flattened.

Physaria ovalifolia

Physaria lepidota

Distribution
from FNA
CO; KS; NE; NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices branched (well-developed); petals usually yellow, rarely white, 6.5-12(-14) mm, usually 1.5 times or less as long as sepals; racemes usually not elongated (subumbellate).
subsp. ovalifolia
1. Caudices usually simple; petals white, (9-)11-15 mm, often 2 times as long as sepals; racemes usually elongated.
subsp. alba
1. Trichomes: rays fused nearly to tips; fruits with deep sinuses, or shallow basally, deep apically.
subsp. lepidota
1. Trichomes: rays fused in proximal 1/2; fruits with sinuses absent or shallow basally, deep apically.
subsp. membranacea
Source FNA vol. 7, p. 655. FNA vol. 7, p. 648.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. ovalifolia subsp. alba, P. ovalifolia subsp. ovalifolia
P. lepidota subsp. lepidota, P. lepidota subsp. membranacea
Synonyms Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia
Name authority (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) Rollins: Brittonia 33: 335, figs. 1, 2. (1981)
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