Physaria ovalifolia |
Physaria gooddingii |
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roundleaf bladderpod |
Goodding's bladderpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate). | ||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate. |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
(proximal usually shortly petiolate, distal sessile); blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed. |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
dense, compact, (elongated in fruit). |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled); petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire). |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(recurved, curved or sigmoid), somewhat expanded apically. |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm; valves pubescent, trichomes spreading, sparsely pubescent inside; ovules 4–6 per ovary; style 3–5 mm. |
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Seeds | flattened. |
flattened. |
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Physaria ovalifolia |
Physaria gooddingii |
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Phenology | Flowering Jun–Sep. | |||||
Habitat | Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests | |||||
Elevation | 1800-2300 m (5900-7500 ft) | |||||
Distribution |
CO; KS; NE; NM; OK; TX
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AZ; NM |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 639. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | Lesquerella gooddingii | ||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |