Physaria ovalifolia |
Physaria didymocarpa |
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roundleaf bladderpod |
common twinpod, double twinpod, twin bladderpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Perennials; caudex branched, (cespitose); densely pubescent, trichomes (often stalked, appressed to wavy and spreading), several-rayed, rays furcate or simple, (slightly to strongly umbonate, nearly smooth to strongly tuberculate). | ||||||||||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
(forming a strong rosette; long-petiolate); blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery). |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute). |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
congested, (elongated in fruit, greatly exceeding leaves). |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals lanceolate to oblong, 6–8 mm, (often keeled); petals spatulate, 10–12 mm. |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(spreading, straight or slightly curved), 8–12 mm. |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy); replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse; ovules (4–)8 per ovary; style 7–9 mm. |
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Seeds | flattened. |
flattened. |
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Physaria ovalifolia |
Physaria didymocarpa |
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Distribution |
CO; KS; NE; NM; OK; TX
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ID; MT; WA; WY; AB; BC
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 633. | ||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | Vesicaria didymocarpa, Coulterina didymocarpa | ||||||||||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||||||||||
Web links |