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roundleaf bladderpod

Arizona bladderpod

Habit Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout).
Stems

few to several from base, erect or outer decumbent, 0.5–2.5 dm.

simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm.

Basal leaves

blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate.

(densely tufted, not rosulate, reflexed in age);

blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute).

Cauline leaves

(proximal shortly petiolate, distal usually sessile);

blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire.

similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm.

Racemes

compact, (± subumbellate to densely corymbiform, elongated or not).

dense, often subcorymbiform.

Flowers

sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically);

petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated).

sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically);

petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect).

Fruiting pedicels

(usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout).

(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm.

Fruits

(sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 4–8(–9) mm.

(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm;

valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth;

ovules 4–10(–16) per ovary;

style (0.5–)1–2(–4) mm (shorter than fruit).

Seeds

flattened.

flattened.

2n

= 10.

Physaria ovalifolia

Physaria arizonica

Phenology Flowering Apr–Jun.
Habitat Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine
Elevation 1000-2200 m (3300-7200 ft)
Distribution
from FNA
CO; KS; NE; NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
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[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices branched (well-developed); petals usually yellow, rarely white, 6.5-12(-14) mm, usually 1.5 times or less as long as sepals; racemes usually not elongated (subumbellate).
subsp. ovalifolia
1. Caudices usually simple; petals white, (9-)11-15 mm, often 2 times as long as sepals; racemes usually elongated.
subsp. alba
Source FNA vol. 7, p. 655. FNA vol. 7, p. 627.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. ovalifolia subsp. alba, P. ovalifolia subsp. ovalifolia
Synonyms Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis
Name authority (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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