Physaria ovalifolia |
Physaria arctica |
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roundleaf bladderpod |
arctic bladderpod |
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Habit | Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo). | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). | ||||
Stems | few to several from base, erect or outer decumbent, 0.5–2.5 dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
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Basal leaves | blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate. |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
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Cauline leaves | (proximal shortly petiolate, distal usually sessile); blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire. |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
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Racemes | compact, (± subumbellate to densely corymbiform, elongated or not). |
loose. |
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Flowers | sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically); petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated). |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
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Fruiting pedicels | (usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout). |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
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Fruits | (sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–16 per ovary; style 4–8(–9) mm. |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
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Seeds | flattened. |
plump. |
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2n | = 60. |
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Physaria ovalifolia |
Physaria arctica |
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Phenology | Flowering May–Aug. | |||||
Habitat | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance | |||||
Elevation | 0-1800 m (0-5900 ft) | |||||
Distribution |
CO; KS; NE; NM; OK; TX
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AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa | ||||
Name authority | (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
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