Physaria oregona |
Physaria hitchcockii |
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Oregon twin-pod |
Hitchcock's bladderpod |
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Habit | Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
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Basal leaves | (petiole slender, usually incised or with broad teeth along petiole); blade obovate, 4–6 cm, margins entire. |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
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Cauline leaves | blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute). |
similar to basal, smaller. |
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Racemes | somewhat loose, (5–15 cm). |
dense. |
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Flowers | sepals oblong, 5–7 mm; petals (lemon yellow), spatulate, 9–12 mm. |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (spreading or ascending, curved, fruits not pendent), 10–20 mm. |
(ascending, straight or slightly curved), 2–6 mm. |
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Fruits | didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading; replum broadly lanceolate, as wide as or wider than fruit, apex acute; ovules 8 per ovary; style 1–2 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
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Seeds | flattened. |
flattened. |
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2n | = 8. |
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Physaria oregona |
Physaria hitchcockii |
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Phenology | Flowering Apr–Jun. | |||||||||
Habitat | Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils | |||||||||
Elevation | 900-1900 m (3000-6200 ft) | |||||||||
Distribution |
ID; OR; WA
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NV; UT |
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Discussion | Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Coulterina oregona, Lesquerella oregona | Lesquerella hitchcockii | ||||||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |