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Oregon twin-pod

Goodding's bladderpod

Habit Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate).
Stems

several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm.

several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm.

Basal leaves

(petiole slender, usually incised or with broad teeth along petiole);

blade obovate, 4–6 cm, margins entire.

blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate.

Cauline leaves

blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute).

(proximal usually shortly petiolate, distal sessile);

blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed.

Racemes

somewhat loose, (5–15 cm).

dense, compact, (elongated in fruit).

Flowers

sepals oblong, 5–7 mm;

petals (lemon yellow), spatulate, 9–12 mm.

sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled);

petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire).

Fruiting pedicels

(spreading or ascending, curved, fruits not pendent), 10–20 mm.

(recurved, curved or sigmoid), somewhat expanded apically.

Fruits

didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open);

valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading;

replum broadly lanceolate, as wide as or wider than fruit, apex acute;

ovules 8 per ovary;

style 1–2 mm.

(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm;

valves pubescent, trichomes spreading, sparsely pubescent inside;

ovules 4–6 per ovary;

style 3–5 mm.

Seeds

flattened.

flattened.

2n

= 8.

Physaria oregona

Physaria gooddingii

Phenology Flowering Apr–Jun. Flowering Jun–Sep.
Habitat Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests
Elevation 900-1900 m (3000-6200 ft) 1800-2300 m (5900-7500 ft)
Distribution
from FNA
ID; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM
[BONAP county map]
Discussion

Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 655. FNA vol. 7, p. 639.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Coulterina oregona, Lesquerella oregona Lesquerella gooddingii
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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