Physaria oregona |
Physaria arizonica |
|
---|---|---|
Oregon twin-pod |
Arizona bladderpod |
|
Habit | Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). |
Stems | several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
Basal leaves | (petiole slender, usually incised or with broad teeth along petiole); blade obovate, 4–6 cm, margins entire. |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
Cauline leaves | blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute). |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
Racemes | somewhat loose, (5–15 cm). |
dense, often subcorymbiform. |
Flowers | sepals oblong, 5–7 mm; petals (lemon yellow), spatulate, 9–12 mm. |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
Fruiting pedicels | (spreading or ascending, curved, fruits not pendent), 10–20 mm. |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
Fruits | didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading; replum broadly lanceolate, as wide as or wider than fruit, apex acute; ovules 8 per ovary; style 1–2 mm. |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
Seeds | flattened. |
flattened. |
2n | = 8. |
= 10. |
Physaria oregona |
Physaria arizonica |
|
Phenology | Flowering Apr–Jun. | Flowering Apr–Jun. |
Habitat | Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine |
Elevation | 900-1900 m (3000-6200 ft) | 1000-2200 m (3300-7200 ft) |
Distribution |
ID; OR; WA
|
AZ; UT
|
Discussion | The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 627. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Coulterina oregona, Lesquerella oregona | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis |
Name authority | S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
Web links |