Physaria oregona |
Physaria acutifolia |
|
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Oregon twin-pod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; caudex simple, (cespitose); (silvery) pubescent throughout, trichomes (stalked), few-rayed, rays furcate or imperfectly so, (tuberculate throughout). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). |
Stems | several from base, erect or somewhat decumbent, (unbranched), 1–3.5 dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
Basal leaves | (petiole slender, usually incised or with broad teeth along petiole); blade obovate, 4–6 cm, margins entire. |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
Cauline leaves | blade oblanceolate or broader, 1.5–2.5 cm, margins entire or sparsely dentate, (apex acute). |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
Racemes | somewhat loose, (5–15 cm). |
loose, (elongated in fruit). |
Flowers | sepals oblong, 5–7 mm; petals (lemon yellow), spatulate, 9–12 mm. |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
Fruiting pedicels | (spreading or ascending, curved, fruits not pendent), 10–20 mm. |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
Fruits | didymous, obreniform, moderately inflated, angustispetate, (8–)10–12(–15) × 10–14(–16) mm, (papery, not keeled, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence, rounded or irregular), loosely pubescent, trichomes spreading; replum broadly lanceolate, as wide as or wider than fruit, apex acute; ovules 8 per ovary; style 1–2 mm. |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
Seeds | flattened. |
(dark brown), flattened, (2–3 mm). |
2n | = 8. |
= 10, 16, 24. |
Physaria oregona |
Physaria acutifolia |
|
Phenology | Flowering Apr–Jun. | Flowering May–Jun(-Jul). |
Habitat | Gravelly banks, stream shores, rocky slopes, dry hillsides, serpentine soils | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities |
Elevation | 900-1900 m (3000-6200 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
ID; OR; WA
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 655. | FNA vol. 7, p. 624. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Coulterina oregona, Lesquerella oregona | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa |
Name authority | S. Watson: Proc. Amer. Acad. Arts 17: 363. (1882) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) |
Web links |