Physaria newberryi |
Physaria tumulosa |
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Newberry twinpod, Newberry's twinpod |
Kodachrome bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex (buried), branched, (forming hard mats); densely pubescent, trichomes several-rayed, rays furcate or bifurcate, (tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect, (unbranched), 0.2–0.3 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(few), similar to cauline. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(petiole not differentiated from blade); blade (somewhat succulent), linear to narrowly oblanceolate, 5–12 mm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, (few-flowered). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (yellowish), elliptic, 3–4.5 mm; petals (erect or, more commonly, arching), spatulate to oblanceolate, 5.8–7 mm, (claw not or weakly differentiated from blade). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(ascending to divaricate-ascending, ± straight), 3.5–6 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(coppery or reddish brown in age), broadly ovoid, slightly inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.8–3 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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Physaria newberryi |
Physaria tumulosa |
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Phenology | Flowering May–Jun. | |||||
Habitat | Barren white knolls surrounded by sagebrush, pinyon pine, and Utah juniper | |||||
Elevation | 1600-1800 m (5200-5900 ft) | |||||
Distribution |
AZ; NM; NV; UT
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UT |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria tumulosa is morphologically similar to 55. P. navajoensis of northeastern Arizona and northwestern New Mexico, and differing very subtly. It has been long treated as an infraspecific taxon of P. hitchcockii; unpublished molecular data do not support that disposition. It is found on knolls of the Winsor Member of the Carmel Formation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 664. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella hitchcockii subsp. tumulosa, Lesquerella tumulosa, P. rubicundula var. tumulosa | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Barneby) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | ||||
Web links |