Physaria newberryi |
Physaria spatulata |
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Newberry twinpod, Newberry's twinpod |
alpine bladderpod, spatula-leaf bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(erect to prostrate, petiole distinct from blade); blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade spatulate, distinctly different from basal. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
moderately dense, (6–20-flowered). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (pale yellow), elliptic, 3.5–5 mm; petals lingulate, 6–9 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(strongly sigmoid), 10–20 mm (2 times longer than fruits). |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed); valves pubescent, trichomes sparse and closely appressed to surface; ovules 4–8 per ovary; style 2.5–6 mm (usually ± equal in length to mature fruit). |
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Seeds | slightly flattened, (ovate). |
plump. |
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Physaria newberryi |
Physaria spatulata |
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Phenology | Flowering May-early Jul. | |||||
Habitat | Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates | |||||
Elevation | 900-2900 m (3000-9500 ft) | |||||
Distribution |
AZ; NM; NV; UT
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MT; ND; NE; SD; WY; AB; SK
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 663. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rydberg) Grady & O’Kane: Novon 17: 190. (2007) | ||||
Web links |