Physaria newberryi |
Physaria scrotiformis |
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Newberry twinpod, Newberry's twinpod |
silver twinpod, west silver bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; (diminutive); caudex simple or branched, (buried, with thatch of persistent leaf bases distally); (appearing silvery gray-green to silvery purple), densely pubescent, trichomes usually 5 or 6 (rarely 7)-rayed, rays bifurcate or incompletely so, (relatively short, stout, umbonate, moderately tuberculate to nearly smooth, lower layer smoother). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
1–5 from base, prostrate to slightly decumbent, (arising laterally, also erect or ascending from tuft of basal leaves, unbranched, purple-green), 0.08–0.3 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(petiole slightly winged); blade oblanceolate, elliptic, or rhombic, (mostly flat, sometimes somewhat folded), 0.6–2.7 cm, (base tapering to petiole), margins entire, (apex rounded to rounded-acute). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(3–7, shortly petiolate or sessile); blade elliptic to oblanceolate, 0.3–0.5 cm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
crowded, (ca. 3–7 fruits). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (greenish yellow), linear-triangular, 3.7–5 mm, (lateral pair subsaccate); petals oblanceolate to narrowly obovate, 4.5–9 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(ascending, straight), 1.8–3.4 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(shortly stipitate, purple or greenish purple in age), slightly didymous, ovoid to obpyriform, 3–5 mm (wider than long, base rounded-obtuse, apex rounded, flattened, or slightly emarginate); valves (inflated, slightly wider than replum), pubescent, trichomes scattered; replum obovate to orbicular-obdeltate, apex rounded, obtuse, or truncate; septum complete or medially small-perforate; ovules 4–6(–8) per ovary; style 2–3.6 mm. |
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Seeds | slightly flattened, (ovate). |
relatively plump, (ovate to suborbicular, usually rounded on one side, ± flat or concave on the other, not mucilaginous when wetted). |
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Physaria newberryi |
Physaria scrotiformis |
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Phenology | Flowering Jun-early Jul. | |||||
Habitat | Tundra areas with islands of Engelmann spruce on Leadville limestone, amidst limestone cobbles and gravel | |||||
Elevation | 3500-3700 m (11500-12100 ft) | |||||
Distribution |
AZ; NM; NV; UT
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CO |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria scrotiformis is known only from La Plata County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 662. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | |||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | O’Kane: Novon 17: 376, fig. 1. (2007) | ||||
Web links |