Physaria newberryi |
Physaria reediana |
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Newberry twinpod, Newberry's twinpod |
alpine bladderpod, reed's twinpod, Rollins' bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple or branched, (covered with persistent leaf bases, loosely cespitose); densely pubescent, trichomes (appressed to ascending, plant appearing shaggy, always appressed on fruits), 4- or 5-rayed, rays furcate or bifurcate, (tuberculate throughout). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
few to several from base, ± erect, (arising laterally, also from within basal leaves), 0.2–0.4 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(erect); blade linear-oblanceolate, 1.2–2.8 cm, (base gradually narrowed to petiole), margins entire. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade linear, similar to basal. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, (often subumbellate, not or barely exceeding basal leaves). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (pale green-yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals lingulate, 6–9 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(ascending, curved), 3–5.5 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
lanceolate in outline, compressed (latiseptate) on margins and at apex, 4–5 mm; valves pubescent, trichomes closely appressed; ovules 8–12 per ovary; style 3.5–4.5 mm (equaling or exceeding length of fruit, curved proximal to stigma). |
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Seeds | slightly flattened, (ovate). |
plump, (oblong). |
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Physaria newberryi |
Physaria reediana |
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Phenology | Flowering May–Jul. | |||||
Habitat | Open areas of grasslands on calcareous soils | |||||
Elevation | 1200-1900 m (3900-6200 ft) | |||||
Distribution |
AZ; NM; NV; UT
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CO; NE; WY
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 660. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Vesicaria alpina, Alyssum alpinum, Lesquerella alpina, Lesquerella alpina var. laevis, Lesquerella condensata var. laevis | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||
Web links |