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Newberry twinpod, Newberry's twinpod

straight bladderpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Perennials; (loose, spreading); caudex simple or branched; densely pubescent, trichomes (subsessile), 4–6-rayed, rays furcate or bifurcate, (moderately tuberculate over arms, less so or smooth over center).
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

few to several from base, ascending or prostrate, (arising laterally, also from within basal leaves, usually unbranched, rarely branched), 0.5–3(–6) dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

blade narrowly oblanceolate or broadly elliptic, 1–7(–12) cm, margins entire or shallowly toothed, sometimes repand, (inner blades usually flattened in age, surfaces often gray-green, scabrous).

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

(usually secund);

blade spatulate or obovate, 1–2.5(–4.5) cm, margins entire or shallowly toothed, (flat or involute).

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

somewhat crowded (to moderately elongated in fruit, exceeding basal leaves).

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

sepals broadly elliptic or oblong, 4–7.5(–9) mm, (median pair thickened apically, cucullate);

petals cuneate or obovate, 7–10(–16) mm, (tapering gradually to broad claw).

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

(often divaricate-spreading and straight, or horizontal and loosely sigmoid, sometimes slightly recurved), 5–15 mm.

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

subglobose to ovoid or ellipsoid, sometimes compressed, (4–)5–7(–9) mm;

valves sparsely pubescent, trichomes appressed or erect, sometimes sparsely pubescent inside;

ovules (8–)12–16(–20) per ovary;

style 2–7 mm.

Seeds

slightly flattened, (ovate).

somewhat flattened.

2n

= 10 + 2, 18, 20, ± 40.

Physaria newberryi

Physaria rectipes

Phenology Flowering May–Jul.
Habitat Sandy soils, limey knolls, rocky hills, clay hillsides, dry ridges, weathered rocks, gravelly outwashes, stony slopes, pinyon-juniper woodlands
Elevation 1500-2600 m (4900-8500 ft)
Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As here circumscribed, Physaria rectipes remains heterogeneous and may represent more than one taxon, even after the recent removal of P. pulvinata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 652. FNA vol. 7, p. 660.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi Lesquerella rectipes
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002)
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