Physaria newberryi |
Physaria pinetorum |
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Newberry twinpod, Newberry's twinpod |
White Mountain bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(petiole tapering to blade); blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(not or loosely overlapping, petiolate or distal sessile); blade spatulate to oblanceolate, 1–4 cm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
crowded, elongated. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate); petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(ascending, curved or sigmoid), 6–12(–20) mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–24 per ovary; style (2–)4–7 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 10. |
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Physaria newberryi |
Physaria pinetorum |
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Phenology | Flowering Apr–Jul. | |||||
Habitat | Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices | |||||
Elevation | 1400-2900 (-3400) m (4600-9500 (-11200) ft) | |||||
Distribution |
AZ; NM; NV; UT
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AZ; NM
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 658. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella pinetorum | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) | ||||
Web links |