Physaria newberryi |
Physaria nelsonii |
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Newberry twinpod, Newberry's twinpod |
Nelson's bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves). |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(absent or few); similar to basal, blade linear. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, (few-flowered). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals (bright yellow), lingulate, 6–8 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(loosely sigmoid), 3–5 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
lanceolate, compressed apically, 2.5–3(–4) mm; valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside; ovules 4–8 per ovary; style 2.5–4 mm. |
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Seeds | slightly flattened, (ovate). |
plump, (oblong). |
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Physaria newberryi |
Physaria nelsonii |
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Phenology | Flowering May-mid Jun. | |||||
Habitat | Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants | |||||
Elevation | 1600-2300 m (5200-7500 ft) | |||||
Distribution |
AZ; NM; NV; UT
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UT; WY |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 652. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |