Physaria newberryi |
Physaria intermedia |
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Newberry twinpod, Newberry's twinpod |
mid-bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(clustered at stem base); blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
compact, (often nearly subumbellate). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate); petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout). |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened); valves sparsely pubescent, trichomes appressed; ovules (8–)12–16(–20) per ovary; style (2–)3–4.5(–5.5) mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 18, 20, 36. |
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Physaria newberryi |
Physaria intermedia |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates | |||||
Elevation | 1600-2400 m (5200-7900 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AZ; NM; UT
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 644. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella alpina var. intermedia, Lesquerella intermedia | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||
Web links |