Physaria newberryi |
Physaria integrifolia |
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Newberry twinpod, Newberry's twinpod |
Snake River or creeping twinpod, Snake River twinpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex usually branched, (rhizomelike, cespitose); densely (silvery) pubescent, trichomes (often stalked, appressed), several-rayed, rays furcate or bifurcate, (umbonate, strongly tuberculate throughout). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, ± erect, exceeding basal rosette by ± 0.5 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(forming a strong rosette; long-petiolate); blade oblanceolate to ovate or orbicular, (1.5–)2–4(–8) cm, (base usually abruptly tapering to petiole), margins entire. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade oblanceolate, 1–2 cm, margin entire, (apex acute). |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
congested, (greatly exceeding leaves). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals often keeled, 6–8 mm; petals spatulate, 8–10 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(spreading, straight or slightly curved), 7–11 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
didymous, highly inflated, 8–22 × 10–25 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), densely pubescent, trichomes appressed; replum linear to oblong, as wide as or wider than fruit; ovules 8 per ovary; style 7–9 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 16. |
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Physaria newberryi |
Physaria integrifolia |
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Phenology | Flowering (May-)Jun–Jul(-Aug). | |||||
Habitat | Calcareous hills and slopes, shale-limestone cliffs, bare steep slopes, red clay banks, shale | |||||
Elevation | 1900-2700 m (6200-8900 ft) | |||||
Distribution |
AZ; NM; NV; UT
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ID; MT; WY |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria integrifolia has traditionally been recognized as a variety of P. didymocarpa, but it is morphologically and ecologically quite distinctive. Variety monticola (no combination has been made at subspecific rank) is not recognized here; it is considered another example, in the genus, of caudices elongating in response to shifting substrates. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 644. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | P. didymocarpa var. integrifolia, P. integrifolia var. monticola | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rollins) Lichvar: Madroño 31: 203. (1984) | ||||
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