Physaria newberryi |
Physaria gooddingii |
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Newberry twinpod, Newberry's twinpod |
Goodding's bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Annuals or biennials; without caudex, (cespitose); densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (ascending or erect), simple or infrequently furcate near base, (long and slender, sometimes with U-shaped notch on one side, smooth or finely tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect (and stout) or outer ones decumbent, (unbranched or branched, stiff and densely foliate, sterile leaf-bearing branches sometimes present), to 4 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade obovate or elliptic, to ca. 3 cm, margins sinuate or shallowly dentate. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal usually shortly petiolate, distal sessile); blade obovate to broadly elliptic, 1–3 cm, margins sinuate or shallowly toothed. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, compact, (elongated in fruit). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals elliptic or narrowly elliptic or oblong, (3.8–)4.5–5.5 mm, (lateral pair cucullate, very convex, median pair tapering to base, thickened apically, cucullate, often slightly keeled); petals cuneate, 6.5–8 mm, (slightly expanded at base, margins sinuate, apex retuse or entire). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(recurved, curved or sigmoid), somewhat expanded apically. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or substipitate), oblong or broadly elliptic, compressed (latiseptate), 5–8 mm; valves pubescent, trichomes spreading, sparsely pubescent inside; ovules 4–6 per ovary; style 3–5 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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Physaria newberryi |
Physaria gooddingii |
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Phenology | Flowering Jun–Sep. | |||||
Habitat | Mountainous areas, open areas in pinyon-juniper and ponderosa pine forests | |||||
Elevation | 1800-2300 m (5900-7500 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AZ; NM |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria gooddingii (found in the mountains of Catron, Sierra, and western Socorro counties, New Mexico, and in Greenlee County, Arizona) is similar to 9. P. aurea (found farther east), but differs in having trichomes with ascending or erect rays (rather than appressed) and fruits that are strongly latiseptate (rather than not, or very little, compressed), a state that is infrequent in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 639. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella gooddingii | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |