Physaria newberryi |
Physaria globosa |
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Newberry twinpod, Newberry's twinpod |
globe bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Biennials or perennials; caudex branched, (± woody); densely pubescent, trichomes (sessile), 3–6-rayed, rays distinct and simple or furcate, (in 2 layers, lower layer umbonate, smooth to finely tuberculate, some often with a U-shaped notch). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect, (arising among leaves of an elongated main axis), to 5 dm (± equal). |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(shortly petiolate); blade obovate to oblanceolate, (1.5–)2.5–5(–6) cm, margins entire, sinuate to shallowly toothed, or pinnatifid. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(sessile or shortly petiolate); blade oblanceolate to oblong, 1.3–3(–4) cm, (base cuneate), margins entire or repand to dentate. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals elliptic or obovate, 2.6–4.1 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate, 3.5–6.5(–7.5) mm, (margins sinuate). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(usually spreading horizontally, straight), 7–14(–21) mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or substipitate); globose, often slightly compressed apically, (1–)2–3 mm; valves sparsely pubescent, sometimes pubescent inside, trichomes spreading, 3–5-rayed; ovules 4 per ovary; style 2–3.5(–4) mm. |
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Seeds | slightly flattened, (ovate). |
flattened or plump, (often outer surface hemispherical, inner surface flattened, or both surfaces rounded). |
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2n | = 14. |
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Physaria newberryi |
Physaria globosa |
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Phenology | Flowering Mar–May. | |||||
Habitat | Open rocky areas, shale at cliff bases, open talus, ledges, open cedar glades | |||||
Elevation | 100-300 m (300-1000 ft) | |||||
Distribution |
AZ; NM; NV; UT
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IN; KY; TN |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria globosa is possibly introduced in Indiana. A report for Ohio was based on a collection by “Jones,” but that specimen cannot be located. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 639. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Vesicaria globosa, Alyssum globosum, Alyssum shortii, Lesquerella globosa | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Desvaux) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |