Physaria newberryi |
Physaria fremontii |
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Newberry twinpod, Newberry's twinpod |
Fremont's bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple; densely pubescent, trichomes (subsessile), 5–7-rayed, rays distinct or slightly fused at base, usually furcate, (roughly tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
few to several from base, prostrate, (arising proximal to a terminal cluster of erect leaves, usually unbranched, slender), 0.5–1(–1.5) dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(petiole slender); blade elliptic to rhombic, 1.5–4(–5) cm, (base gradually tapering to petiole), margins usually entire, rarely dentate. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade narrowly obovate to oblanceolate, 5–15 mm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
(relatively short), rather loose. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ± elliptic, 4–6 mm; petals ovate, 6–8 mm, (claw slightly expanded at base). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(usually secund, recurved), 5–8 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(pendent), globose, subglobose, or slightly obcompressed, slightly or not inflated, 3–6 mm, (rigid, apex usually beaked); valves pubescent throughout; ovules 8–12 per ovary; style 1–2 mm, (pubescent). |
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Seeds | slightly flattened, (ovate). |
slightly flattened, (suborbicular). |
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Physaria newberryi |
Physaria fremontii |
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Phenology | Flowering May–Jun. | |||||
Habitat | Calcareous gravel, loose whitish rubble, limestone pavement, rocky calcareous ridges | |||||
Elevation | 2100-2800 m (6900-9200 ft) | |||||
Distribution |
AZ; NM; NV; UT
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WY |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria fremontii is known from the area of the Wind River Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 638. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella fremontii | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |