Physaria newberryi |
Physaria filiformis |
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Newberry twinpod, Newberry's twinpod |
limestone glade or Missouri bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Annuals; with a fine taproot; densely pubescent, trichomes (sessile), usually 4-rayed, rays forked, rarely simple or tripartite, (finely tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, (slender), erect or outer decumbent, (usually branched, branches filiform, bud clusters of growing plants drooping), to 2.5 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade orbicular to broadly spatulate, 1–2.4 cm, margins entire or sinuate. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal often petiolate, distal sessile); similar to basal, blade spatulate to oblanceolate or (distal) linear, (base cuneate), margins entire or sinuate. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
loose. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals oblong or elliptic, 2.5–4.6 mm, (median pair slightly thickened apically); petals (pale yellow), spatulate to obovate, 5–9 mm, (apex ± emarginate). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(usually divaricate-ascending, straight or slightly curved), 7–11 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or shortly stipitate), globose, not inflated, 3–4 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 3–5 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 14. |
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Physaria newberryi |
Physaria filiformis |
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Phenology | Flowering Apr–May. | |||||
Habitat | Limestone, dolomite, and shale, sparsely vegetated or barren areas, cedar glades, old pastures, along roadsides | |||||
Elevation | 200-300 m (700-1000 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AL; AR; MO |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 637. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella filiformis | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |