Physaria newberryi |
Physaria engelmannii |
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Newberry twinpod, Newberry's twinpod |
Engelmann's bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple or branched, (woody, aerial); densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays simple or furcate, distinct or fused at base, (asymmetrical with deep notch on one side, often with a U-shaped gap between 2 of the rays, umbonate, strongly tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
few to several from base, erect, (usually unbranched), (1.5–)2.5–4(–6) dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade elliptic to obovate, 2–6.5 cm, margins entire, sinuate, or remotely toothed, (surfaces occasionally sparsely pubescent). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal often petiolate, distal sessile or subsessile); blade oblanceolate to linear, 1–4 cm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, (subumbellate). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ovate or elliptic, 5.5–10 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate to elliptic, 8–14 mm, (sometimes with distinct claw, often retuse). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(ascending), relatively short. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(shortly stipitate), ± globose or ellipsoid, not or slightly inflated, 5–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20 per ovary; style 3.5–5 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 12, 24, 36. |
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Physaria newberryi |
Physaria engelmannii |
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Phenology | Flowering Apr–May. | |||||
Habitat | Limestone prairies, rocky ridges, pebbly shores, thin caliche soils, limestone outcrops | |||||
Elevation | 150-400 m (500-1300 ft) | |||||
Distribution |
AZ; NM; NV; UT
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OK; TX |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 635. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Vesicaria engelmannii, Alyssum engelmannii, Lesquerella engelmannii, Vesicaria engelmannii var. elatior, Vesicaria pulchella | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |